Numbers beside nodes are ML bootstrap proportions ≥70 (above branches) and Bayesian posterior probabilities ≥90 (under branches). All rRNA+COI haplotypes contained the 12S and 16S sequences, and all COI+rRNA haplotypes contained the COI sequence. Triangles indicate nodes that are not supported in the Bayesian analysis. Asterisk represents Q. robertingeri which is thought to be the synonym of Q. boulengeri (Che et al., 2009).